Hands are always tricky. Getting the thumbs facing the right way, drawing the angles correctly, especially the bend of the wrist...
So, when John came by with a cup of tea, I got him to pose. He had to sit on the floor, to get the angle right. The extra flourishes, in the 'roaring' department, were his own idea (he really threw himself into the role!).
I did a very quick sketch, capturing the shapes but imagining his fingers as being more dumpy and paw-like:
I then transferred the pose to my lion. This is a detail from the final rough of the spread. Much better: